was less interested in prehistory and in simpler, nonliterate societies. Yet the roots of inequality in the modern world lie far back in prehistory. Hence Toynbee did not pose Yali's question, nor did he come to grips with what I see as history's broadest pattern. Other available books on world history similarly tend to focus on advanced literate Eurasian civilizations of the last 5,000 years; they have a very brief treatment of pre-Columbian Native American civilizations, and an even briefer discussion of the rest of the world except for its recent interactions with Eurasian civilizations. Since Toynbee's attempt, worldwide syntheses of historical causation have fallen into disfavor among most historians, as posing an apparently intractable problem. Specialists from several disciplines have provided global syntheses of their subjects. Especially useful contributions have been made by ecological geographers, cultural anthropologists, biologists studying plant and animal domestication, and scholars concerned with the impact of infectious diseases on history. These studies have called attention to parts of the puzzle, but they provide only pieces of the needed broad synthesis that has been missing. Thus, there is no generally accepted answer to Yali's question. On the one hand, the proximate explanations are clear: some peoples developed guns, germs, steel, and other factors conferring political and economic power before others did; and some peoples never developed these power factors at all. On the other hand, the ultimate explanations—for example, why bronze tools appeared early in parts of Eurasia, late and only locally in the New World, and never in Aboriginal Australia—remain unclear. Our present lack of such ultimate explanations leaves a big intellectual gap, since the broadest pattern of history thus remains unexplained. Much more serious, though, is the moral gap left unfilled. It is perfectly obvious to everyone, whether an overt racist or not, that different peoples have fared differently in history. The modern United States is a European- YALI'SQUESTION • 25 molded society, occupying lands conquered from Native Americans and incorporating the descendants of millions of sub-Saharan black Africans brought to America as slaves. Modern Europe is not a society molded by sub-Saharan black Africans who brought millions of Native Americans as slaves. These results are completely lopsided: it was not the case that 51 percent of the Americas, Australia, and Africa was conquered by Europeans, while 49 percent of Europe was conquered by Native Americans, Aboriginal Australians, or Africans. The whole modern world has been shaped by lopsided outcomes. Hence they must have inexorable explanations, ones more basic than mere details concerning who happened to win some battle or develop some invention on one occasion a few thousand years ago. It seems logical to suppose that history's pattern reflects innate differences among people themselves. Of course, we're taught that it's not polite to say so in public. We read of technical studies claiming to demonstrate inborn differences, and we also read rebuttals claiming that those studies suffer from technical flaws. We see in our daily lives that some of the conquered peoples continue to form an underclass, centuries after the conquests or slave imports took place. We're told that this too is to be attributed not to any biological shortcomings but to social disadvantages and limited opportunities. Nevertheless, we have to wonder. We keep seeing all those glaring, persistent differences in peoples' status. We're assured that the seemingly transparent biological explanation for the world's inequalities as of a.d. 1500 is wrong, but we're not told what the correct explanation is. Until we have some convincing, detailed, agreed-upon explanation for the broad pattern of history, most people will continue to suspect that the racist biological explanation is correct after all. That seems to me the strongest argument for writing this book. authors are regularly asked by journalists to summarize a long book in one sentence. For this book, here is such a sentence: 'History followed different courses for different peoples because of differences among peoples' environments, not because of biological differences among peoples themselves.' Naturally, the notion that environmental geography and biogeography influenced societal development is an old idea. Nowadays, though, the 26 •PROLOGUE view is not held in esteem by historians; it is considered wrong or simplistic, or it is caricatured as environmental determinism and dismissed (ha as did Cambridge Prof of history Martin Daunton!), or else the whole subject of trying to understand worldwide differences is shelved as too difficult. Yet geography obviously has some effect on history; the open question concerns how much effect, and whether geography can account for history's broad pattern. The time is now ripe for a fresh look at these questions, because of new information from scientific disciplines seemingly remote from human history. Those disciplines include, above all, genetics, molecular biology, and biogeography as applied to crops and their wild ancestors; the same disciplines plus behavioral ecology, as applied to domestic animals and their wild ancestors; molecular biology of human germs and related germs of animals; epidemiology of human diseases; human genetics; linguistics; archaeological studies on all continents and major islands; and studies of the histories of technology, writing, and political organization. This diversity of disciplines poses problems for would-be authors of a book aimed at answering Yali's question. The author must possess a range of expertise spanning the above disciplines, so that relevant advances can be synthesized. The history and prehistory of each continent must be similarly synthesized. The book's subject matter is history, but the approach is that of science—in particular, that of historical sciences such as evolutionary biology and geology. The author must understand from firsthand experience a range of human societies, from hunter-gatherer societies to modern space-age civilizations. These requirements seem at first to demand a multi-author work. Yet that approach would be doomed from the outset, because the essence of the problem is to develop a unified synthesis. That consideration dictates single authorship, despite all the difficulties that it poses. Inevitably, that single author will have to sweat copiously in order to assimilate material from many disciplines, and will require guidance from many colleagues. My background had led me to several of these disciplines even before Yali put his question to me in 1972. My mother is a teacher and linguist; my father, a physician specializing in the genetics of childhood diseases. Because of my father's example, I went through school expecting to become a physician. I had also become a fanatical bird-watcher by the age of seven. It was thus an easy step, in my last undergraduate year at university, to shift from my initial goal of medicine to the goal of biological YALI'SQUESTION • 27 research. However, throughout my school and undergraduate years, my training was mainly in languages, history, and writing. Even after deciding to obtain a Ph.D. in physiology, I nearly dropped out of science during my first year of graduate school to become a linguist. Since completing my Ph.D. in 1961, I have divided my scientific research efforts between two fields: molecular physiology on the one hand, evolutionary biology and biogeography on the other hand. As an unforeseen bonus for the purposes of this book, evolutionary biology is a historical science forced to use methods different from those of the laboratory sciences. That experience has made the difficulties in devising a scientific approach to human history familiar to me. Living in Europe from 1958 to 1962, among European friends whose lives had been brutally traumatized by 20th-century European history, made me start to think more seriously about how chains of causes operate in history's unfolding. For the last 33 years my fieldwork as an evolutionary biologist has brought me into close contact with a wide range of human societies. My specialty is bird evolution, which I have studied in South America, southern Africa, Indonesia, Australia, and especially New Guinea. Through living with