formation of coral atolls, which suggested, not coincidentally, that atolls could not form in less than a million years- the first hint of his long-standing attachment to the extreme antiquity of earthly processes. In 1836, aged twenty- seven, he returned home after being away for five years and two days. He never left England again.

One thing Darwin didn’t do on the voyage was propound the theory (or even a theory) of evolution. For a start, evolution as a concept was already decades old by the 1830s. Darwin’s own grandfather, Erasmus, had paid tribute to evolutionary principles in a poem of inspired mediocrity called “The Temple of Nature” years before Charles was even born. It wasn’t until the younger Darwin was back in England and read Thomas Malthus’s Essay on the Principle of Population (which proposed that increases in food supply could never keep up with population growth for mathematical reasons) that the idea began to percolate through his mind that life is a perpetual struggle and that natural selection was the means by which some species prospered while others failed. Specifically what Darwin saw was that all organisms competed for resources, and those that had some innate advantage would prosper and pass on that advantage to their offspring. By such means would species continuously improve.

It seems an awfully simple idea-it is an awfully simple idea-but it explained a great deal, and Darwin was prepared to devote his life to it. “How stupid of me not to have thought of it!” T. H. Huxley cried upon reading On the Origin of Species. It is a view that has been echoed ever since.

Interestingly, Darwin didn’t use the phrase “survival of the fittest” in any of his work (though he did express his admiration for it). The expression was coined five years after the publication of On the Origin of Species by Herbert Spencer in Principles of Biology in 1864. Nor did he employ the word evolution in print until the sixth edition of Origin (by which time its use had become too widespread to resist), preferring instead “descent with modification.” Nor, above all, were his conclusions in any way inspired by his noticing, during his time in the Galapagos Islands, an interesting diversity in the beaks of finches. The story as conventionally told (or at least as frequently remembered by many of us) is that Darwin, while traveling from island to island, noticed that the finches’ beaks on each island were marvelously adapted for exploiting local resources-that on one island beaks were sturdy and short and good for cracking nuts, while on the next island beaks were perhaps long and thin and well suited for winkling food out of crevices-and it was this that set him to thinking that perhaps the birds had not been created this way, but had in a sense created themselves.

In fact, the birds had created themselves, but it wasn’t Darwin who noticed it. At the time of the Beagle voyage, Darwin was fresh out of college and not yet an accomplished naturalist and so failed to see that the Galapagos birds were all of a type. It was his friend the ornithologist John Gould who realized that what Darwin had found was lots of finches with different talents. Unfortunately, in his inexperience Darwin had not noted which birds came from which islands. (He had made a similar error with tortoises.) It took years to sort the muddles out.

Because of these oversights, and the need to sort through crates and crates of other Beagle specimens, it wasn’t until 1842, six years after his return to England, that Darwin finally began to sketch out the rudiments of his new theory. These he expanded into a 230-page “sketch” two years later. And then he did an extraordinary thing: he put his notes away and for the next decade and a half busied himself with other matters. He fathered ten children, devoted nearly eight years to writing an exhaustive opus on barnacles (“I hate a barnacle as no man ever did before,” he sighed, understandably, upon the work’s conclusion), and fell prey to strange disorders that left him chronically listless, faint, and “flurried,” as he put it. The symptoms nearly always included a terrible nausea and generally also incorporated palpitations, migraines, exhaustion, trembling, spots before the eyes, shortness of breath, “swimming of the head,” and, not surprisingly, depression.

The cause of the illness has never been established, but the most romantic and perhaps likely of the many suggested possibilities is that he suffered from Chagas’s disease, a lingering tropical malady that he could have acquired from the bite of a Benchuga bug in South America. A more prosaic explanation is that his condition was psychosomatic. In either case, the misery was not. Often he could work for no more than twenty minutes at a stretch, sometimes not that.

Much of the rest of his time was devoted to a series of increasingly desperate treatments-icy plunge baths, dousings in vinegar, draping himself with “electric chains” that subjected him to small jolts of current. He became something of a hermit, seldom leaving his home in Kent, Down House. One of his first acts upon moving to the house was to erect a mirror outside his study window so that he could identify, and if necessary avoid, callers.

Darwin kept his theory to himself because he well knew the storm it would cause. In 1844, the year he locked his notes away, a book called Vestiges of the Natural History of Creation roused much of the thinking world to fury by suggesting that humans might have evolved from lesser primates without the assistance of a divine creator. Anticipating the outcry, the author had taken careful steps to conceal his identity, which he kept a secret from even his closest friends for the next forty years. Some wondered if Darwin himself might be the author. Others suspected Prince Albert. In fact, the author was a successful and generally unassuming Scottish publisher named Robert Chambers whose reluctance to reveal himself had a practical dimension as well as a personal one: his firm was a leading publisher of Bibles. Vestiges was warmly blasted from pulpits throughout Britain and far beyond, but also attracted a good deal of more scholarly ire. The Edinburgh Review devoted nearly an entire issue-eighty-five pages-to pulling it to pieces. Even T. H. Huxley, a believer in evolution, attacked the book with some venom, unaware that the author was a friend.[42]

Darwin’s manuscript might have remained locked away till his death but for an alarming blow that arrived from the Far East in the early summer of 1858 in the form of a packet containing a friendly letter from a young naturalist named Alfred Russel Wallace and the draft of a paper, On the Tendency of Varieties to Depart Indefinitely from the Original Type, outlining a theory of natural selection that was uncannily similar to Darwin’s secret jottings. Even some of the phrasing echoed Darwin’s own. “I never saw a more striking coincidence,” Darwin reflected in dismay. “If Wallace had my manuscript sketch written out in 1842, he could not have made a better short abstract.”

Wallace didn’t drop into Darwin’s life quite as unexpectedly as is sometimes suggested. The two were already corresponding, and Wallace had more than once generously sent Darwin specimens that he thought might be of interest. In the process of these exchanges Darwin had discreetly warned Wallace that he regarded the subject of species creation as his own territory. “This summer will make the 20th year (!) since I opened my first note-book, on the question of how amp; in what way do species amp; varieties differ from each other,” he had written to Wallace some time earlier. “I am now preparing my work for publication,” he added, even though he wasn’t really.

In any case, Wallace failed to grasp what Darwin was trying to tell him, and of course he could have no idea that his own theory was so nearly identical to one that Darwin had been evolving, as it were, for two decades.

Darwin was placed in an agonizing quandary. If he rushed into print to preserve his priority, he would be taking advantage of an innocent tip-off from a distant admirer. But if he stepped aside, as gentlemanly conduct arguably required, he would lose credit for a theory that he had independently propounded. Wallace’s theory was, by Wallace’s own admission, the result of a flash of insight; Darwin’s was the product of years of careful, plodding, methodical thought. It was all crushingly unfair.

To compound his misery, Darwin’s youngest son, also named Charles, had contracted scarlet fever and was critically ill. At the height of the crisis, on June 28, the child died. Despite the distraction of his son’s illness, Darwin found time to dash off letters to his friends Charles Lyell and Joseph Hooker, offering to step aside but noting that to do so would mean that all his work, “whatever it may amount to, will be smashed.” Lyell and Hooker came up with the compromise solution of presenting a summary of Darwin’s and Wallace’s ideas together. The venue they settled on was a meeting of the Linnaean Society, which at the time was struggling to find its way back into fashion as a seat of scientific eminence. On July 1, 1858, Darwin’s and Wallace’s theory was unveiled to the world. Darwin himself was not present. On the day of the meeting, he and his wife were burying their son.

The Darwin-Wallace presentation was one of seven that evening-one of the others was on the flora of Angola-and if the thirty or so people in the audience had any idea that they were witnessing the scientific highlight

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